(click for larger image)
Reconstruction of O. reburrus by M. Collins. The precise arrangement of the anteriormost region remains somewhat conjectural.
Halkieriids are Cambrian animals that looked like slugs in scale mail; often when they died their scales, called sclerites, dissociated and scattered, and their sclerites represent a significant component of the small shelly fauna of the early Cambrian. They typically had their front and back ends capped with shells that resembled those we see in bivalve brachiopods. Wiwaxiids were also sluglike, but sported very prominent, long sclerites, and lacked the anterior and posterior shells; their exact position in the evolutionary tree has bounced about quite a bit, but some argument has made that they belong in the annelid ancestry, and that their sclerites are homologous to the bristly setae of worms. One simplistic picture of their relationship to modern forms was that the halkieriids expanded their shells and shed their scales to become molluscs, while the wiwaxiids minimized their armor to emphasize flexibility and became more wormlike. (Note that that is a very crude summary; relationships of these Cambrian groups to modern clades are extremely contentious. There’s a more accurate description of the relationships below.)
Now a new fossil has been found, Orthozanclus reburrus that unites the two into a larger clade, the halwaxiids. Like the halkieriids, it has an anterior shell (but not a posterior one), and like the wiwaxiids, it has long spiky sclerites. In some ways, this simplifies the relationships; it unites some problematic organisms into a single branch on the tree. The question now becomes where that branch is located—whether the halwaxiids belong in a separate phylum that split off from the lophophorate family tree after the molluscs, or whether the halwaxiids are a sister group to the molluscs.
